# Biomedical Sciences

**Source**: https://catalog.ucsd.edu/faculty/BIOM.html
**Parent**: https://catalog.ucsd.edu/front/courses.html

[ [program](https://catalog.ucsd.edu/curric/BIOM.html) | [courses](https://catalog.ucsd.edu/courses/BIOM.html) ]

*All courses, faculty listings, and curricular and degree requirements described herein are subject to change or deletion without notice.*

#### Professors

Ruben Abagyan, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Cristian Achim, MD, PhD, *Psychiatry and Neuropathology (Adjunct)*

Susan Ackerman, PhD, *Cellular and Molecular Medicine*

Joseph Adams, PhD, *Pharmacology*

Rommie Amaro, PhD, *Chemistry and Biochemistry*

Corina Antal, PhD, *Pharmacology*

Radha Ayyagari, PhD, *Radiology/Ophthalmology*

Maksim Bazhenov, PhD, *Medicine*

Bogdan Bintu, PhD, *Cellular and Molecular Medicine*

Gerry R. Boss, MD, *Medicine*

Nunzio Bottini, MD, PhD, *Medicine*

Joan Heller Brown, PhD, *Pharmacology (Chair)*

Laurence L. Brunton, PhD, *Pharmacology/Medicine (Emeritus)*

Jack Bui, MD, PhD, *Pathology*

Michael Burkart, PhD, Professor, *Chemistry and Biochemistry*

Geoffrey Chang, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

John Chang, MD, *Medicine*

Fabian Rivera-Chávez, PhD, *Pediatrics*

Ju Chen, PhD, *Medicine (In Residence)*

David Cheresh, PhD, *Pathology*

Neil Chi, MD, PhD, *Medicine*

Jerold J. M. Chun, PhD, *Pharmacology (Adjunct)*

Don W. Cleveland, PhD, *Cellular and Molecular Medicine (Chair)/Medicine*

Nicole Coufal, MD, PhD, *Pediatrics (Adjunct)*

Leslie A. Crews, PhD, *Medicine*

Michael Croft, PhD, *Medicine*

Shane Crotty, PhD, *Medicine (Adjunct)*

Antonio De Maio, PhD, *Surgery*

Edward Dennis, PhD, *Pharmacology/Chemistry and Biochemistry*

Arshad Desai, PhD, *Cellular and Molecular Medicine*

Neal Devaraj, PhD, *Chemistry and Biochemistry*

Jack E. Dixon, PhD, *Pharmacology/Cellular and Molecular Medicine/Chemistry and Biochemistry (Dean, Scientific Affairs)*

Daniel Donoghue, PhD, *Chemistry and Biochemistry*

Pieter Dorrestein, PhD, *Pharmacy/Chemistry and Biochemistry*

Steven F. Dowdy, PhD, *Cellular and Molecular Medicine*

Tal Einav, PhD, *La Jolla Institute of Immunology*

Mark H. Ellisman, PhD, *Neurosciences*

Adam Engler, PhD, *Bioengineering/UC San Diego Stem Cell Institute*

Peter Ernst, DVM, PhD, *Pathology*

Jeffrey Esko, PhD, *Cellular and Molecular Medicine*

Ronald M. Evans, PhD, *Biology (Adjunct)*

Sylvia Evans, PhD, *Pharmacy/Pharmacology*

Xi Fang, PhD, *Medicine*

Gen-Sheng Feng, PhD, *Pathology*

Cole Ferguson, MD, PhD, *Pathology*

Fleur M. Ferguson, PhD, *Chemistry and Biochemistry/Skaggs School of Pharmacy and Pharmaceutical Sciences*

Susan Ferro-Novick, PhD, *Cellular and Molecular Medicine*

Kelly Frazer, PhD, *Pediatrics*

Theodore Friedmann, MD, *Pediatrics*

Frank Funari, PhD, *Pathology*

Pascal Gagneux, PhD, *Cellular and Molecular Medicine (Adjunct)*

Richard Gallo, MD, PhD, *Medicine*

William Gerwick, PhD, *Oceanography/Skaggs School of Pharmacy and Pharmaceutical Sciences*

Partho Ghosh, PhD, *Chemistry and Biochemistry*

Pradipta Ghosh, MD, *Medicine*

Mark Ginsberg, MD, *Medicine*

Christopher K. Glass, MD, PhD, *Cellular and Molecular Medicine/Medicine*

Joseph G. Gleeson, MD, *Neurosciences*

Ananda Goldrath, PhD, *Molecular Biology*

Steven Gonias, MD, PhD, *Pathology (Chair)*

Philip Gordts, PhD, *Department of Medicine*

John C. Guatelli, PhD, *Medicine*

Shiri Gur-Cohen, PhD, *Medicine*

Asa Gustafsson, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Silvio Gutkind, PhD, *Pharmacology*

Gabriel Haddad, MD, *Pediatrics (Chair)/Neurosciences*

Bruce Hamilton, PhD, *Cellular and Molecular Medicine/Medicine*

Tracy Handel, PhD, *Skaggs School of* *Pharmacy and Pharmaceutical Sciences*

Stephen Hedrick, PhD, *Cellular and Molecular Medicine*

Sven Heinz, PhD, *Medicine*

Robert Hevner, MD, PhD, *Pathology*

Michael C. Hogan, PhD, *Medicine (Adjunct)*

Vivian Hook, PhD, *Skaggs School of* *Pharmacy and Pharmaceutical Sciences*

Anthony Hunter, PhD, *Molecular Biology*

Trey Ideker, PhD, *Medicine*

Catriona Jamieson, MD, PhD, *Medicine*

Yishi Jin, PhD, *Cellular and Molecular Medicine/Biological Sciences*

Michael Karin, PhD, *Pharmacology*

Alexander Kauffman, PhD, *Reproductive Medicine*

Dan Kaufman, MD, PhD, *Medicine*

John Kelsoe, MD, *Psychiatry*

David Kleinfeld, PhD, *Physics*

Rob Knight, PhD, *Pediatrics*

Richard Kolodner, PhD, *Cellular and Molecular Medicine*

Elizabeth A. Komives, PhD, *Chemistry and Biochemistry*

Mitchell Kronenberg, PhD, *Molecular Biology*

Mark Lawson, PhD, *Reproductive Medicine*

Hyam L. Leffert, MD, *Pharmacology*

Andres Leschziner, PhD, *Cellular and Molecular Medicine*

Willis Li, PhD, *Medicine*

George Liu, MD, PhD, *Pediatrics*

James McKerrow, MD, PhD, *Dean, Skaggs School of Pharmacy and Pharmaceutical Sciences*

Hiruy Meharena, PhD, *Neurobiology*

Jill Mesirov, PhD, *Medicine*

Yury Miller, MD, PhD, *Medicine*

Bradley Moore, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Alysson Muotri, PhD, *Pediatrics/Cellular and Molecular Medicine*

Cornelis Murre, PhD, *Molecular Biology*

Samuel Myers, PhD, *La Jolla Institute for Immunology*

Alexandra Newton, PhD, *Pharmacology*

Victor Nizet, MD, *Pediatrics and Skaggs School of Pharmacy and Pharmaceutical Sciences*

Peter Novick, PhD, *Cellular and Molecular Medicine*

Karen Oegema, PhD, *Cellular and Molecular Medicine*

Abraham Palmer, PhD, *Psychiatry*

Mana Parast, MD, PhD, *Pathology*

Hemal Patel, PhD, *Anesthesiology*

Renate B. Pilz, MD, *Medicine (In Residence)*

Frank L. Powell Jr., PhD, *Medicine*

Oswald Quehenberger, PhD, *Medicine*

Manuela Raffatellu, MD, *Pediatrics*

Tariq Rana, PhD, *Pediatrics (Adjunct)*

Anjana Rao, PhD, *Pharmacology (Adjunct)*

Samara L. Reck-Peterson, PhD, *Cellular and Molecular Medicine*

Bing Ren, PhD, *Cellular and Molecular Medicine*

Michael G. Rosenfeld, MD, *Medicine*

Robert Ross, MD, *Medicine (In Residence)*

Subhojit Roy, MD, PhD, *Pathology*

Alan Saltiel, PhD, *Medicine*

Guy Salvesen, PhD, *Pathology (Adjunct)*

Maike Sander, MD, *Pediatrics/Cellular and Molecular Medicine*

Richard Scheuermann, PhD, *Pathology*

David Schlaepfer, PhD, *Reproductive Medicine*

Geert Schmid-Schoenbein, PhD, *Bioengineering*

Stephen Schoenberger, PhD, *Medicine*

Johannes Schoeneberg, PhD, *Pharmacology*

Nicholas Schork, PhD, *Psychiatry and Biostatistics*

Jonathan Sebat, PhD, *Psychiatry/Cellular and Molecular Medicine*

Jianhua Shao, MD, PhD, *Pediatrics*

Sanford Shattil, MD, *Medicine*

Reuben Shaw, PhD, *Molecular Biology*

Aleem Siddiqui, PhD, *Medicine*

Dionicio Siegel, PhD, *Skaggs School of Pharmacy and Pharmaceutical* *Sciences*

Christina Sigurdson, DVM, PhD, *Pathology*

Monique Smith, PhD, *Neurobiology*

Evan Snyder, MD, PhD, *Pediatrics (Associate Physician)*

Francesca Soncin, PhD, *Pathology*

Deborah Spector, PhD, *Cellular and Molecular Medicine*

Nicole Steinmetz, PhD, *NanoEngineering*

Ina Stelzer, PhD, *Pathology*

Xin Sun, PhD, *Pediatrics*

Roger Sunahara, PhD, *Pharmacology*

Palmer W. Taylor, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Susan S. Taylor, PhD, *Chemistry and Biochemistry, Pharmacology*

David Traver, PhD*, Cellular and Molecular Medicine*

JoAnn Trejo, PhD, *Pharmacology*

Emily Troemel, PhD, *Cell and Developmental Biology*

Robert H. Tukey, PhD, *Pharmacology/Chemistry and Biochemistry*

Ajit P. Varki, MD, *Medicine*

Judith A. Varner, PhD, *Pathology*

Francisco Villarreal, MD, PhD, *Medicine (Adjunct)*

Dong Wang, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Emily Wang, PhD, *Pathology*

Jean Wang, PhD, *Medicine*

Nicholas J. G. Webster, PhD, *Medicine (In Residence)*

Miles Wilkinson, PhD, *Reproductive Medicine*

Karl Willert, PhD, *Cellular and Molecular Medicine*

Elizabeth Winzeler, PhD, *Pediatrics*

Joseph L. Witzum, MD, *Medicine*

Tony L. Yaksh, PhD, *Anesthesiology/Pharmacology*

Jing Yang, PhD, *Pharmacology/Pediatrics*

Gene Yeo, PhD, *Cellular and Molecular Medicine*

Wei Ying, PhD, *Medicine*

Maurizio Zanetti, MD, *Medicine (Emeritus)*

Karsten Zengler*,* PhD, *Pediatrics*

Dong-Er Zhang, PhD, *Pathology/Biological Sciences*

Jin Zhang, PhD, *Pharmacology*

Kun Zhang, PhD, *Bioengineering*

Binhai Zheng, PhD, *Neurosciences*

Huilin Zhou, PhD, *Cellular and Molecular Medicine*

Yimin Zou, PhD, *Neurobiology*

Elina Zuniga, PhD, *Molecular Biology*

#### Associate Professors

Fotis Asimakopoulos, MD, PhD, *Medicine*

Ferhat Ay, PhD, *Pediatrics (Adjunct)*

Carlo Ballatore, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Rafael Bejar, MD, PhD, *Internal Medicine*

Alejandro (Alex) Chavez, PhD, *Pediatrics*

Stephanie Cherqui, PhD, *Pediatrics (In Residence*)

Cosimo Commisso, PhD, *Surgery*

Kevin Corbett, PhD, *Cellular and Molecular Medicine*

Ben Croker, PhD, *Pediatrics*

Kamil Godula, PhD, *Chemistry and Biochemistry*

Thomas Hnasko, PhD, *Neurosciences*

Lilia Iakoucheva, PhD, *Psychiatry*

Juliana Idoyaga, PhD, *Pharmacology and Molecular Biology*

Mohit Jain, MD, PhD, *Medicine, Pharmacology*

William Joiner, PhD, *Pharmacology*

Irina Kufareva, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Nathan Lewis, PhD, *Pediatrics*

Li-Fan Lu, PhD, *Molecular Biology*

Ayako Makino, PhD, *Medicine*

David McCulley, MD, *Pediatrics*

Jerome Mertens, PhD, *Neurosciences*

Anthony Molina, PhD, *Medicine*

Gerald P. Morris, MD, PhD, *Pathology*

Larissa Podust, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Christopher Putnam, PhD, *Medicine*

Zoran Radic, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Robert Rissman, PhD, *Neuroscience*

Simon Schenk, PhD, *Orthopaedic Surgery*

George Sen, PhD, *Medicine/Cellular and Molecular Medicine*

Farah Sheikh, PhD, *Medicine (Adjunct)*

Dwayne Stupack, PhD, *Reproductive Medicine*

Pandurangan Vijayanand, MD, PhD, *Medicine*

Jill Wildonger, PhD, *Pediatrics*

Peter Zage, MD, PhD, *Pediatrics*

Sheng Zhong, PhD*, Bioengineering*

#### Assistant Professors

Ludmil Alexandrov, PhD, *Cellular and Molecular Medicine*

Christopher Benner, PhD, *Medicine*

Zea Borok, PhD, *Medicine*

Aaron Carlin, MD, PhD, *Medicine*

Yuan Chen, PhD, *Surgery*

Hiuting Chu, PhD, *Pathology*

Heidi Cook-Andersen, MD, PhD, *Reproductive Medicine and Biological Sciences*

Richard Daneman, PhD, *Pharmacology and Neuroscience*

Matthew Daugherty, PhD, *Molecular Biology*

Emma Farley, PhD, *Medicine, Molecular Biology*

Stephanie Fraley, PhD, *Bioengineering*

Kyle Gaulton, PhD, *Pediatrics*

Jack A. Gilbert, PhD, *Pediatrics*

David Gonzalez, PhD, *Pharmacology*

Alon Goren, PhD, *Medicine*

Melissa Gymrek, PhD, *Medicine*

Matt Hangauer, PhD, *Dermatology*

Ryan Hibbs, PhD, *Neurobiology*

Wendy Huang, PhD, *Cellular and Molecular Medicine*

Enfu Hui, PhD, *Cell and Developmental Biology*

Louise Laurent, MD, PhD, *Obstetrics, Gynecology, and Reproductive Sciences*

Amanda L. Lewis, PhD, *Obstetrics, Gynecology, and Reproductive Sciences*

Byungkook Lim, PhD, *Neurobiology*

Amit Majithia, MD, *Medicine and Pediatrics*

Prashant Mali, PhD, *Bioengineering*

Karen Mestan, MD, *Pediatrics*

Eran Mukamel, PhD, *Cognitive Science*

Anthony O’Donoghue, PhD, *Skaggs School of Pharmacy and Pharmaceutical Sciences*

Erica Ollmann Saphire, PhD, *Medicine (Adjunct)*

Andrew Sharabi, MD, PhD, *Clinical Radiation Medicine and Applied Science*

Robert Signer, PhD, *Medicine*

Tatum Simonson, PhD, *Medicine*

Scott Sternson, PhD, *Neurosciences*

Bryan Sun, MD, PhD, *Dermatology*

Francesca Telese, PhD, *Medicine*

Elizabeth Villa, PhD, *Chemistry and Biochemistry*

Karl Wahlin, PhD, *Opthalmology*

Samuel R. Ward, PhD, *Orthopaedic Surgery and Radiology*

Daniela Weiskopf, PhD, *Medicine (Adjunct)*

Amir Zarrinpar, MD, PhD, *Medicine*